Bibliographic information and links to the fulltext of peer-reviewed publications and preprints of our lab appear here. Thesis documents are linked to on the Resources page. Following green open-access policies, PDFs are to be used for scientific purposes only. Full list also available on Pubmed and Google Scholar via the buttons to the right.
2018
Domdei N*, Domdei L*, Reiniger J, Linden M, Holz FG, Roorda A, Harmening WM (2018)
Ultra-high contrast retinal display system for single photoreceptor psychophysics.
Biomedical Optics Express, 9: 157-172 [doi:10.1364/BOE.9.000157] [PDF]
2017
2016
2015
2014
Harmening WM*, Tuten WS*, Roorda A, Sincich LC (2014) Mapping the perceptual grain of the human retina. The Journal of Neuroscience, 34(16):5667-5677 [doi:10.1523/JNEUROSCI.5191-13.2014] [PDF]
2012
before 2012
(* denotes equal contribution)
Reiniger JL, Lobecke AC, Sabesan R, Bach M, Verbakel F, de Brabander J, Holz FG, Berendschot TTJM, Harmening WM (2019) Habitual higher order aberrations affect Landolt but not Vernier acuity. Journal of Vision, 19(5): 1-15 [doi:10.1167/19.5.11] [PDF]
Meyer JH, Larsen PF, Strack C, Harmening WM, Krohne TU, Holz FG, Schmitz-Valckenberg S (2019) Optical coherence tomography angiography (OCT-A) in an animal model of laser-induced choroidal neovascularization. Experimental Eye Research, 184: 162-171 [doi:10.1016/j.exer.2019.04.002] [PDF]
Domdei N, Linden M, Reiniger JL, Holz FG, Harmening WM (2019) Eye tracking-based estimation and compensation of chromatic offsets for multi-wavelength retinal microstimulation with foveal cone precision. Biomedical Optics Express, 10:4126-4141. [doi:10.1364/BOE.10.004126] [PDF]
2019
Harmening WM, Sincich LC (2019) Adaptive Optics for Photoreceptor-Targeted Psychophysics. High resolution Imaging in Microscopy and Ophthalmology, Springer, Cham. pp 359-375 [doi:10.1007/978-3-030-16638-0_17] [PDF]
Harmening WM, Göbbels K, Wagner H (2007) Vernier acuity in barn owls.
Vision Research, 47(7): 1020-1026 [doi:10.1016/j.visres.2007.01.005] [PDF]
Harmening WM, Vobig MA, Walter P, Wagner H (2007) Ocular aberrations in barn owl eyes. Vision Research, 47(23): 2934-2942 [doi:10.1016/j.visres.2007.08.001] [PDF]
Ohayon S, Harmening WM, Wagner H, Rivlin E (2008) Through a barn owl's eyes: interactions between scene content and visual attention. Biological Cybernetics,
98(2): 115-132 [doi:10.1007/s00422-007-0199-4] [PDF]
Harmening WM (2009) A case of quasi-infinite visual acuity and illusionary size.
Perception, 38(5): 781-783 [doi:10.1068/p6398
] [PDF]
Harmening WM, Nikolay P, Orlowski J, Wagner H (2009) Spatial contrast sensitivity and grating acuity of barn owls. Journal of Vision, 9(7): 1-12 [doi:10.1167/9.7.13] [PDF]
van der Willigen RF*, Harmening WM*, Vossen S, Wagner H (2010) Disparity sensitivity in man and owl: psychophysical evidence for equivalent perception of shape-from-stereo. Journal of Vision, 10(1): 1-11 [doi:10.1167/10.1.10] [PDF]
Harmening WM, Orlowski J, Ben-Shahar O, Wagner H (2011) Overt attention towards oriented objects in free viewing barn owls. Proceedings of the National Academy of Sciences, 108(20): 8461-8466 [doi:10.1073/pnas.1101582108] [PDF]
Harmening WM, Wagner H (2011) From optics to attention: Visual perception in the barn owl. Journal of Comparative Physiology A, 197(11): 1031-42
[doi:10.1007/s00359-011-0664-3] [PDF]
Orlowski J, Harmening WM, Wagner H (2012) Night vision in barn owls: visual acuity and contrast sensitivity under dark adaptation. Journal of Vision, 12(4):1-8
[doi:10.1167/12.13.4] [PDF]
Scriba M, Harmening WM, Mettke-Hofman C, Vyssotski A, Roulin A, Wagner H, Rattenborg N (2012) Evaluation of two minimally-invasive techniques for electroencephalogram recording in wild or freely behaving animals. Journal of Comparative Physiology A, 199(3):183-189 [doi:10.1007/s00359-012-0779-1] [PDF]
Harmening WM, Tiruveedhula P, Roorda A, Sincich LC (2012) Measurement and correction of transverse chromatic offsets for multi-wavelength retinal microscopy.
Biomedical Optics Express, 3(9): 2066-2077 [doi:10.1364/BOE.3.002066] [PDF]
Bruce KS, Harmening WM, Langston BR, Tuten WS, Roorda A, Sincich LC (2015) Normal perceptual sensitivity arising from weakly reflective cone photoreceptors.
Investigative Ophthalmology & Visual Science, 56(8):4431-4438
[doi:10.1167/iovs.15-16547] [PDF]
Fang PP, Harmening WM, Müller PL, Lindner M, Krohne TU, Holz FG (2015) Technical principles of OCT angiography. Der Ophthalmologe, 113(1):6-13
[doi:10.1007/s00347-015-0184-6] [PDF]
Müller PL, Müller S, Gliem M, Küpper K, Holz FG, Harmening WM*, Issa P* (2016) Perception of Haidinger brushes in macular disease depends on macular pigment density and visual acuity. Investigative Ophthalmology & Visual Science, 57(3): 1448-1456 [doi:10.1167/iovs.15-19004] [PDF]
Meyer J, Fang PP, Krohne TU, Harmening WM, Holz FG, Schmitz-Valckenberg S (2016) Optical coherence tomography angiography (OCT-A) in rats. Der Ophthalmologe,
114(2): 140-147 [doi:10.1007/s00347-016-0309-6] [PDF]
Pfau M, Lindner M, Fleckenstein M, Finger RP, Rubin G,  Harmening WM, Morales MU, Holz FG, Schmitz-Valckenberg S (2016) Test–retest reliability of scotopic and mesopic fundus-controlled perimetry using a modified MAIA in normal eyes. Ophthalmologica,
237: 42-54 [doi:10.1159/000453079] [PDF]
Domdei N, Reiniger JL, Pfau M, Charbel Issa P, Holz FG, Harmening WM (2016) Histology in the living eye: non-invasive microscopic structure and function analysis of the retina with adaptive optics. Der Ophthalmologe, 114(3): 206-214
[doi:10.1007/s00347-016-0411-9] [PDF]
Sincich LC, Sabesan R, Tuten WS, Roorda A, Harmening WM (2016) Functional Imaging of Cone Photoreceptors. In: Baraas R, Marshall J, Kremers J (eds.) Human Color Vision.
Volume 5 of the Springer Series in Vision Research. pp 71-104
[doi:10.1007/978-3-319-44978-4_3] [PDF]
Reiniger JL, Domdei N, Holz FG, Harmening WM (2017) Technical principles of adaptive optics in ophthalmology. Der Ophthalmologe, 114(3): 198-205
[doi:10.1007/s00347-017-0440-z] [PDF]
Harmening WM (2017) Adaptive optics for Ophthalmology.
Der Ophthalmologe, 114(3): 196-197 [doi:10.1007/s00347-016-0413-7] [PDF]
Ratnam K, Domdei N, Harmening WM*, Roorda A* (2017) Benefits of retinal image motion at the limits of spatial vision. Journal of Vision, 17(1): 1-11 [doi:10.1167/17.1.30] [PDF]
Reiniger JL*, Domdei N*, Pfau M, Müller PL, Holz FG, Harmening WM (2017) Potential of adaptive optics for the diagnostic evaluation of hereditary retinal diseases.
Klinische Monatsblätter der Augenheilkunde, 234: 311-319. [doi:10.1055/s-0043-100631] [PDF]
Pfau M, Lindner M, Mueller, PL, Birtel J, Finger RP, Harmening WM, Fleckenstein M, Holz FG, Schmitz-Valckenberg S (2017) Effective dynamic range and retest-reliability of two-color scotopic fundus-controlled perimetry using the S-MAIA in patients with macular diseases. Investigative Ophthalmology & Visual Science, 58(6): 158-167 [doi:10.1167/iovs.17-21454] [PDF]
Harmening WM (2017) [Contrast sensitivity and visual acuity in animals] Kontrastempfindlichkeit und Sehschärfe bei Tieren. Der Ophthalmologe, 114(11): 986-996 [doi:10.1007/s00347-017-0561-4] [PDF]
Tuten WS*, Harmening WM*, Sabesan R, Roorda A, Sincich LC (2017) Spatiochromatic interactions between individual cone photoreceptors in the human retina.
The Journal of Neuroscience, 37(39): 9498-9509 [doi:10.1523/JNEUROSCI.0529-17.2017] [PDF]
Birtel J, Harmening WM, Krohne TU, Holz FG, Issa P, Herrmann P (2017) Retinal injury following laser pointer exposure—a systematic review and case series.
Deutsches Ärzteblatt International, 114: 831-837 [doi:10.3238/arztebl.2017.0831] [PDF]
Fang PP, Domdei N, Herrmann P, Schmitz-Valckenberg S, Holz FG, Harmening WM, Krohne TU (2017) Minimal optical coherence tomography B-scan density for reliable detection of intra- and subretinal fluid in patients with macular diseases. Retina, 39(1): 150-156 [doi:10.1097/IAE.0000000000001918] [PDF]
2021
Neitz A, Jiang X, Kuchenbecker JA, Domdei N, Harmening WM, Yan H, Yeonan-Kim, Patterson SS., Neitz M, Neitz J, Coates DR, Sabesan R (2020) Effect of cone spectral topography on chromatic detection sensitivity. Journal of the Optical Society of America A, 37(4):A244-A254. [doi:10.1364/JOSAA.382384] [PDF]
2020
Reiniger JL, Domdei N, Holz FG, Harmening WM (2021) Human gaze is systematically offset from the center of cone topography. Current Biology, 31(18): P4188-4193
[
doi:10.1016/j.cub.2021.07.005] [PDF]
Tuten WS, Harmening WM (2021) Foveal vision. Current Biology, 31(11):R701-R703. [doi:10.1016/j.cub.2021.03.097] [PDF]
Domdei N, Reiniger JL, Holz FG, Harmening WM (2021) The relationship between visual sensitivity and eccentricity, cone density and outer segment length in the human foveola. Investigative Ophthalmology & Visual Science, 62 (9):1-16
[doi:10.1167/iovs.62.9.31] [PDF]
2022
Birtel J, Gliem M, Harmening WM, Holz FG (2022) Ophthalmic phenotyping: Imaging. In: Clinical Ophthalmic Genetics and Genomics, pp: 53-62
[
doi:10.1016/B978-0-12-813944-8.00038-X] [PDF]
Hofmann J*, Domdei L*, Jainta S, Harmening WM (2022) Assessment of binocular fixational eye movements including cyclotorsion with split-field scanning laser ophthalmoscopy. Journal of Vision, 22(10):5, 1-13
[
doi:10.1167/jov.22.10.5] [PDF]
Ameln J, Reiniger JL, Hess K, Holz, FG, Harmening WM (2022) Supernormal foveal photoreceptor density in Alport Syndrome: A case report.
Eur. J. Ophthal.
[doi:10.1177/11206721221093197] [PDF]
Wynne N, Cava JA, Gaffney M, Heitkotter H, Scheidt A, Reiniger JL, Grieshop J, Yang K, Harmening WM, Cooper RF, Carroll J (2022) Intergrader agreement of foveal cone topography using adaptive optics scanning light ophthalmoscopy. Biomed. Opt. Expr., 13(8), pp:4445-4454. [doi:10.1364/BOE.460821] [PDF]
2023
Fassbender K, Baumert PM, Wintergerst MWM, Terheyden JH, Aslan B, Harmening WM, Ettinger U (2023) GABAergic involvement in selective attention. Journal of Cognitive Neuroscience, 35:6, pp.1-14 [doi:10.1162/jocn_a_01989] [PDF]
Click here to download our PubMed referenced directory of retinal pathologies studied with adaptive optics, in XLS or PDF format (last update Oct. 2023, compiled by Niklas Domdei, Maximilian Pfau and Julius Ameln):


Domdei N, Ameln J, Gutnikov A, Witten JL, Holz FG, Wahl S, Harmening WM (2023) Cone density is correlated to outer segment length and retinal thickness in the human foveola. Investigative Ophthalmology & Visual Science, 64(15):11, 1-11
[doi:10.1167/iovs.64.15.11] [PDF] [Suppl.] [Data]
2024
Ameln J*, Saßmannshausen M*, von der Emde L*, Carmichael-Martins A, Holz FG, Ach T, Harmening WM (2024) Assessment of local sensitivity in incomplete retinal-pigment-epithelium and outer retinal atrophy (iRORA) lesions in intermediate age-related macular degeneration (iAMD). BMJ Open Ophthalmology, 9:e001638
[doi:10.1136/bmjophth-2024-001638] [PDF]
Baumert PM, Fassbender K, Wintergerst MWM, Terheyden JH, Aslan B, Foulsham T, Harmening WM, Ettinger U (2024) Effects of Lorazepam on saccadic eye movements - Evidence from prosaccade and free viewing tasks. Psychopharmacology
[doi:10.1007/s00213-024-06672-z] [PDF]
Mecê P, Gocho K, Harmening WM, Rossi E, Young L (2024) Editorial: Advances in ophthalmic optical imaging: new developments, clinical applications and perspectives. Frontiers Ophthalmology, [doi:10.3389/fopht.2024.1496015] [PDF].
Witten JL, Lukyanova V, Harmening WM (2024) Sub-cone visual resolution by active, adaptive sampling in the human foveola. eLife, 2024 Jun 28v2
[doi:10.7554/eLife.98648.3] [PDF] [Supplement] [Data]
2025
Nghiem T-A E*, Reiniger JL*, Dufour O, Harmening WM*, da Silveira RA* (2022) Fixational eye movements as active sensation for high visual acuity. PNAS. [in press]
Saßmannshausen M*, Ameln J*, von der Emde L*, Holz FG, Ach T, Harmening WM (2024) Evaluation of retinal sensitivity in complete retinal-pigment-epithelium and outer retinal atrophy (cRORA) lesions in intermediate age-related macular degeneration (iAMD) by high-resolution microperimetry. Journal of Clinical Medicine, 13(24), 7785
[doi:10.3390/jcm13247785] [PDF]